Activities of key metabolic enzymes and their respective mRNA transcript levels in amphipods exposed to an experimental warming scenario

DOI

Lake Baikal is inhabited by more than 300 endemic amphipod species, which are narrowly adapted to certain thermal niches due to the high interspecific competition. In contrast, the surrounding freshwater fauna is commonly represented by species with large-scale distribution and high phenotypic thermal plasticity. Here, we investigated the thermal plasticity of the energy metabolism in two closely-related endemic amphipod species from Lake Baikal (Eulimnogammarus verrucosus; stenothermal and Eulimnogammarus cyaneus; eurythermal) and the ubiquitous Holarctic amphipod Gammarus lacustris (eurythermal) by exposure to a summer warming scenario (6-23.6 °C; 0.8 °C d-1). In concert with routine metabolic rates, activities of key metabolic enzymes increased strongly with temperature up to 15 °C in E. verrucosus, whereupon they leveled off (except for lactate dehydrogenase). In contrast, exponential increases were seen in E. cyaneus and G. lacustris throughout the thermal trial (Q10-values: 1.6-3.7). Cytochrome-c-oxidase, lactate dehydrogenase, and 3-hydroxyacyl-CoA dehydrogenase activities were found to be higher in G. lacustris than in E. cyaneus, especially at the highest experimental temperature (23.6 °C). Decreasing gene expression levels revealed some thermal compensation in E. cyaneus but not in G. lacustris. In all species, shifts in enzyme activities favored glycolytic energy generation in the warmth. The congruent temperature-dependencies of enzyme activities and routine metabolism in E. verrucosus indicate a strong feedback-regulation of enzymatic activities by whole organism responses. The species-specific thermal reaction norms reflect the different ecological niches, including the spatial distribution, distinct thermal behavior such as temperature-dependent migration, movement activity, and mating season.

Identifier
DOI https://doi.org/10.1594/PANGAEA.931524
Related Identifier IsSupplementTo https://doi.org/10.1038/s41598-021-83748-2
Related Identifier References https://doi.org/10.1594/PANGAEA.845942
Metadata Access https://ws.pangaea.de/oai/provider?verb=GetRecord&metadataPrefix=datacite4&identifier=oai:pangaea.de:doi:10.1594/PANGAEA.931524
Provenance
Creator Jakob, Lena ORCID logo; Vereshchagina, Kseniya P; Tillmann, Anette; Rivarola-Duarte, Lorena; Axenov-Gribanov, Denis V ORCID logo; Bedulina, Daria S ORCID logo; Gurkov, Anton N ORCID logo; Drozdova, Polina; Timofeyev, Maxim A ORCID logo; Stadler, Peter F ORCID logo; Luckenbach, Till ORCID logo; Pörtner, Hans-Otto ORCID logo; Sartoris, Franz-Josef ORCID logo; Lucassen, Magnus ORCID logo
Publisher PANGAEA
Publication Year 2021
Rights Creative Commons Attribution-NoDerivs 4.0 International; https://creativecommons.org/licenses/by-nd/4.0/
OpenAccess true
Representation
Resource Type Dataset
Format application/vnd.openxmlformats-officedocument.spreadsheetml.sheet
Size 34.7 kBytes
Discipline Earth System Research
Spatial Coverage (105.069W, 51.903S, 105.072E, 51.921N)